Ewens, W. J. 2000 "The Mathematical Foundations of Population Genetics" "Evolutionary Genetics" Singh, R.S. and Krimbas, C.B. (editors) Cambridge University Press, pp. 24-40. See esp. pg. 32 where Ewens criticizes Haldane's concept of the cost of natural selection as misguided due to inappropriate mathematical modeling. Ewens also points out here that Mayr's well known arguments against "bean-bag genetics" were directed specifically against Haldane's substitution cost arguments (see Ewens, 1993 and Mayr 1963).

Burger, R. 2000 "The Mathematical Theory of Selection, Recombination, and Mutation" John Wiley & Sons, Ltd., pg. 110 - 111
"For well taken criticisms, in particular, concerning the substitutional load, see Ewens (1979)."
This quote refers to Ewens arguments against the importance of the substitutional load for Kimura's neutral theory.

Brock, James P. 2000. "The Evolution of Adaptive Systems" Academic Press, pg. 450

Hughes, Austin L 1999. "Adaptive Evolution of Genes and Genomes" Oxford University Press, pp. 40 -41

Burt, Austin 1997 [A review of Kimura's collected papers] Quarterly Review Biology 72, pg. 73

Phelps, F.M. 1995  "A Model for the Evolution of the Genome - The Effect of Stochasticity on Genetic Loads" IMA Journal of Mathematics Applied in Medicine and Biology 12: 1-11

Ewens, W. J. 1993 "Beanbag Genetics and After" "Human Population Genetics" Majumder, P.P. (editor) Plenum Press, pp. 7 - 29

Phelps, F.M. 1991  "Multicomponent Rank Selection as an Alternative to Haldane's Dilemma" IMA Journal of Mathematics Applied in Medicine and Biology 8: 57-72

Phelps, F.M. 1991  "A Unifying Model for the Substitutional Genetic Load" IMA Journal of Mathematics Applied in Medicine and Biology 8: 31-56

Gillespie, J.H. 1991 "The Causes of Molecular Evolution" Oxford University Press, pp. 269-272

Wallace, Bruce 1991 "Fifty Years of Genetic Load - An Odyssey" Cornell University Press
See particularly Chapter 5, Dilemmas and Options; Chapter 6, Hard and Soft Selection; Chapter 8, Self-Culling and the Persistance of Populations; and Chapter 9, Summarizing Remarks for issues related to Haldane's Cost of Natural Selection.

Dronamraju, K. R. (editor) 1988 "Haldane and Modern Biology", pp. 165 - 178 - "The Cost of Evolution and Genetic Loads" deals exclusively with these issues.

Wills, C. 1981 "Genetic Variability", pp. 22 - 35

Hartl, D. L. 1980 "Priciples of Population Genetics"  pp. 377 - 378

Ewens, W. J. 1979 "Mathematical Population Genetics"  pp. 66-69, pp. 252-255

Spiess, E. 1977 "Genes in Populations" pp. 619-620

Ewens, W. J. 1973 Comments on Dr. Kimura's Paper Genetics Supplement 73: 36-38

Kimura, M. and Ohta, T. 1973 Mutation and Evolution at the Molecular Level Genetics Supplement 73 pp. 19-35

Ewens, W. J. 1973 Comments on Nei's Letter on Substitutional Loads American Naturalist 107:462-463

Nei, M. 1973 Ewens on the Substitutional Load American Naturalist 107:459-462

Ewens, W. J. 1972 Concepts of Substitutional Load in Finite Populations Theoretical Population Biology 3:153-161

Ewens, W. J. 1972 The Substitutional Load in a Finite Population American Naturalist 106:273-282

Felsenstein, J. 1971 On the Biological Significance of the Cost of Gene Substitution. American Naturalist 105: 1-11

Felsenstein, J. 1971 The Substitution Load in a Finite Population Heredity 28: 57-69

Moran, P. A. P. 1970"Haldane's Dilemma" and the Rate of Evolution Annals of Human Genetics 33:245-249

Ewens, W. J. 1970 Remarks on the Substitutional Load Theoretical Population Biology 1:129-139

O'Donald, P. 1969 "Haldane's Dilemma" and the Rate of Natural Selection Nature 221:815-816

Kimura, M. and Maruyama T. 1969 The Substitutional Load in a Finite Population Heredity 24:101-114

Kimura, M. and Crow, J. F. 1969 Natural Selection and Gene Substitution Genet. Res.

Sved, J. A. 1968 Possible Rates of Gene Substitution in Evolution" The American Naturalist 102:283-293

Maynard-Smith, J. 1968 "Haldane's Dilemma" and the Rate of Evolution Nature 219:1114-1116

 Kimura, M. 1968 Evolutionary Rate at the Molecular Level  Nature 217:624-626

 Crow, J. F. 1968 The Cost of Evolution and Genetic Loads, In Haldane and Modern Biology, John Hopkins Press, Baltimore

 Feller, W. 1967 On Fitness and the Cost of Natural Selection Genetical Research 9:1-15

 Feller, W. 1966 On the Influence of Natural Selection on Population Size Proceedings of the National Academy of Science 55:733-738

Brues, A. M. 196? Genetic Loads and Its Varieties Science 164:1130-1136

Brues, A. M. 1964 The Cost of Evolution Vs. the Cost of Not Evolving Evolution 18:379-383

Van Valen, L. 1963 American Naturalist 47:185

Mayr, E. 1963 "Animal Species and Evolution" Harvard University Press

Dodson, E.O. 1962 Note on the Cost of Natural Selection The American Naturalist 46:123-126

Haldane, J. B. S. 1960 More Precise Expressions for the Cost of Natural Selection Journal of Genetics 57:351-360

Haldane, J. B. S. 1957 The Cost of Natural Selection Journal of Genetics 55:511-524

Please email me (Robert Williams) at rwms@gate.net with comments, suggestions, or questions.